Fox FamilyTreeDNA Project Website - Results

Updated 1/15/2013

Project results as of the above date can be summarized as follows:

Haplogroup R-S26/L1 - The Null439 Foxes:

The null439 Fox group of four family lines has now has established three well-documented distant cousin relationships. The Philadelphia (Fox1) clan has a third cousin, twice removed, relationship (14179 and 37645) with four mutations in 37 markers separated by only 10 transmission events. (A fifth mutation has been observed at DYF406S1 in the last 30 markers of the 67 marker series.) The other two members of this group, 25525 and 25549, are brother and nephew to 14179 and match him exactly at 25 markers. The British (Fox2) clan (16564 and 60400) consists of seventh cousins who are separated by 14 transmission events and have only one mutation in 37 markers. There is a third member of this family tested outside the project at 17 markers who matches both of them at those 17 markers. The South Carolina (Fox3) clan has fourth cousins once removed (25481 and 52944) and second cousins once removed (25481 and 96656) with no mutations in a combined 14 transmission events. 25481 and 96656 are compared at 67 markers while 52944 has been tested at 37 markers. (Note: FTDNA’s 14-14-16-16 result for 25481 at DYS 464 is still listed in this project's Y-Results table but, in reality, should be changed to 14-16-16-16 based on the results of further testing at two other labs.) The three branches have thus exhibited markedly different mutation rates. A common ancestor for the three null439 Fox families, some 10 to 15 generations back, is definitely a possibility."

A fourth null 439 Fox family (Fox4) has now been identified with the addition of 157124 to the project in October 2009.  The odds are good that he is in the same null 439 Fox line but of a separate branch that may share a common Fox ancestor in England back in the 1500's. He is a descendant of Thomas Dudley Fox – who was born in Vermont between 1802 and 1806 and moved to Ontario, Canada.   See the write-up in Project Background for further details.

Another addition to the group is 48348, who traces back to a George Clark of Lycoming County, PA, but is a close match to the Fox3 null439 group (a 66 for 67 marker match with 25481.)  It is known that George Clark was adopted by the Clark family. A possible paper trail to the null439 Fox family via a Mathias Fox connection is being investigated. Mathias moved west from Lycoming County to Clearfield County around 1850. Could Mathias have been a descendant of John Fox, another non-Quaker member of the Plymouth Friends who sailed to Philadelphia in 1686?

205722 has a very deep relationship to this group.  He is also null at DYS 439 and has tested as Haplogroup R-L1.  He is a descendant of Johan Caspar Voss, born in Germany in 1677.  205722 is a genetic distance of about 20 with the other Foxes in this grouping at 67 markers.  Tested also by, he has a total of 76 markers tested.

Six members of this group (excluding 205722 from the group) have been tested at 67 markers.  Aside from 157124 (the Dudley Fox descendant) there was only one mutation in the last 30 markers; that being for 14179 who has 11 repeats at DYF406S1 rather than 10 repeats for all others.  36645, also in the Fox1 group, has been found to be 10 repeats at DYF406S1 so the ancestral 67 marker haplotype for all three of these null439 Fox groups is well defined.  The Fox1 group has undergone the greatest number of mutations but the overall relationship between Fox1, Fox2 and Fox3 is well established.

The markers which have varied for this group are:

DYS 391 = 11 for all but Fox2b, who is 12:  modal = 11
DYS 458 = 17 for all but Fox1a, 1b and 1c, who are 16 and Clark, who is 18:  modal = 17
DYS 447 = 25 for all but Fox1 group, who are 26: modal = 25
DYS 576 = 18 for Fox1a, all Fox3 and Clark, 19 for Fox1d, and 17 for all Fox2:  modal = 18
DYS 570 = 17 for all but Fox1a, who is 16:  modal = 17
CDYa,b = 38, 38 for all but Fox1d, who is 38, 39:  modal = 38, 38
DYS 406S1 = 10 for all but Fox1a, who is 11: modal = 10
The R-S26/L1 modal equals the null439 Fox modal for all of these markers.

The Fox4 family line is represented by 157124, Fox4a, tested at 67 markers. There are some significant differences from the other null439 Foxes but, all in all, the similarities outweigh the differences, which may be summarized for Fox4a as:

Same as null439 Fox Modal at DYS 391, DYS 458, DYS 447, DYS 570 and DYS 406S1

DYS 464a = 15 where above Fox modal = 14 and R-S26 modal = 15.
576 = 17 where above Foxes are 17, 18 and 19 and  null 439 Fox modal is 18.
CDYa,b = 40, 40 where above Fox modal is 38, 38.
DYS 413a,b = 23, 25  where above Foxes and R1b modal = 23, 23.
DYS  534 = 16  where the above Foxes are 534 = 14 and R1b modal is 534 = 15.
DYS  617 = 13  where the above Foxes and the R1b modal  = 12.

The genetic distance of Fox4a from the other null439 Foxes is 10 or 11 but this is basically only on 6 markers and might represent as few as 6 single event changes, with some being multiple steps.  FTDNA’s TiP analysis indicates that the probability of a common ancestor for Fox4a and Fox2a within 20 generations (roughly 500 years) is roughly 90%.  It thus seems possible that a common ancestor did live in England in the 1400’s or 1500’s but tying this down will be difficult.  Some possibilities for going back in time are indicated in the Project Background section.  Certainly, getting more of Thomas Dudley Fox’s descendants tested will be important.

Significant progress has been made in identifying null439 haplotypes in other surnames and classifying them into subclusters.  There are two main subclusters separated by mutations at DYS459b, a stable marker on the palindrome section of the Y-chromosome where a repair mechanism is at work during meiosis (a process occurring during conception.) A Null439 Website has been set up at FTDNA, making it possible for anyone with a null result at DYS 439 to join simply by clicking on a link on his personal webpage.
Neal Fox administers the null439 Project at FTDNA and has set up the Leo Little Memorial Fund to provide for additional tests on certain null493s at key markers that help define clusters.  As a result of Neal's efforts, three main clusters and several subclusters have now been identified..

Project member 14179, one of the original null439 group, was SNP tested by Ethnoancestry and found to be M269+ and S21+ (same as U106). The former is indicative of Haplogroup R1b1b2 (very common) and the latter is a more recently identified SNP, which some propose may indicate a North Sea heritage. He was also the first person to be typed S26+ by EthnoAncestry, a brand new SNP indicative of null439 status.  The 2009 ISOGG Haplogroup designation is R1b1b2a1a1c.  FTDNA named this SNP L1, in honor of Leo Little, and now include it in their Deep Clade R testing.  They call the Haplogroup R1b1b2a1a3.  Since none of the Foxes have been tested L1 by FTDNA, they still show the Haplogroup as R1b1b2 on the Y-Results tables on this Fox Project website. 

Project members 14179 and 25481 have been found negative for the L132 SNP, which appears to split the R-L1/S26 subclade, but will require further verification. This potential SNP was discovered by Neal Fox (co-administrator of the Fox Project and administrator of the Null439 Project at FTDNA) by looking at results from 23andMe, where all null 439's in his cluster 1 had a no-call at this location.  Results from Geno 2.0 may help define more subclades of R-S26/L1.

14179 and 25481 were also tested by DNA-Fingerprint at several markers in the palindrome section of the Y-chromosome and the test results tend to confirm an event within the null439 (S26+) subclade where the backup copy overwrites the other copy during meiosis making large changes to such markers in a single event. This is known as gene conversion or a recLOH event (recombination loss of heterozygosity).   [DNA-Fingerprint results for 14179 were: 464X = 14c,16c,16g, 16g; DYF371 = 10c,12t,13c,14c ; DYF399 = 22, 24, 25.1 ; DYS434 = 9.  For 25481, they were: 464X = 14c,16c,16g, 16g; DYF399 = 22, 24, 26.1; DYS434 = 9.]  14179 and 25481 have now both been tested at over 80 some markers with only 6 single step mutations being recorded.

The Vanfossen Group:

The Van Fossen group now has 17 reported results due to efforts of Clay Fox, the project leader. Eleven of these have a common ancestor in Arnold Van Vossen, a Dutch Mennonite immigrant to Germantown, PA, in the 1700s. Their 37 marker results confirm a Fox/Vanfossen relationship predicted from a handwritten note on a will. Four (24049, 24972, 25721 and 27152) trace back to Oxford Township, Chester County, Pennsylvania, with a Peter Vanfossen as the common ancestor, and another six (38640, 43080, 95996, 95997, 117169 and 171369) trace back to Arnold Van Vossen directly.

There are another five who are not Arnold Van descendants.  Four are residents of the Netherlands, are not a match for the above ten.  Two (35689 and 36279) trace back to Abraham Van Vossen of Bathmen, Netherlands, and are a 25 for 25 match.  116167 and 117168 descend from Jacob Van Vossen 1660 and Willem Van Vossen 1880 of the Netherlands.  Like an American Van Fossen (155575) they do not match the rest of the group.

There are deviations within the group of ten at DYS 459a, DYS 456 and CDYa,b but the ancestral haplotype for Arnold Van Vossen is now pretty well established.  The modal result is DYS 459a = 9, DYS 456 = 18 and CDYa,b = 35,37. The first three of this group (24049, 24972 and 25721) represent father, son and grandson and, interestingly, show the most variation.  They are distinguished from the others by a value of 36 repeats at CDYb. The latest Van Fossen to join the project, 171369, is exactly on the modal.

These are exciting results and a splinter VanFossen Project has been set up at World Families. We will continue to monitor their progress.

Haplogroup R1b - Southern USA Foxes

Enough Southern US Fox descendants have been identified in the R-M269 Haplogroup that they have been grouped together in the Y-DNA results tabulation as "Southern US R-M269." Many members of this Southern R-M269 Group had suspected a connection to Henry Fox and Anne West of Virginia but we now have a definitive set of five results (common ancestor being Henry Fox’s grandson Henry Fox, III,) that appear to define this family line and distinguish it from other Fox families.  We would still like to take the common ancestors back all the way to Henry Fox and Anne West and even beyond.

Henry Fox/ Anne West Descendants

Joseph Steadman and others have exhaustively documented the family line of gentleman Henry Fox, who came to Virginia, married Anne West, a grand niece of Lord De La Warr, and left many descendants in the United States.  He was the son of John Fox, a sea captain who also settled in Virginia in 1661, and this line has been traced back to a Henry Fox (1521) who married a Hawes of Missenden and possibly to a William Fox (1497-1559) of Missenden, Buckinghamshire, who lived at Stewkley Manor.  A William Vaux, descended from a Norman Invader named Robert de Vaux, is known to have inherited Stewkley Manor by marriage in 1424. If a Fox/Vaux connection is substantiated, this would carry the line back to 1066.

We now have six well-documented members of this line in the project. For the original three of these, the most recent common ancestor is Henry Fox (1768-1852), who married Sarah Harrell.   Two are fourth cousins (i.e. 10 transmission events separating them) whose common ancestor is Henry Fox (1768-1852) who married Sarah Harrell. These two are 48275 and 85202 and they match exactly at 37 markers. 48275 and 99137 are second cousins and they match at 25 markers.The haplotype of Henry Fox (1768-1852) can thus be inferred to be the same as theirs.

108898 and 117867, second cousins, take the connection back two generations more to Henry Fox, III (1698-1770), grandson of Henry Fox(1650-1714) and Anne West, who married Mary Goodwyn. They differ from 48275 and85202 only at fast moving marker CDYa, where they have 35 rather than 36repeats. So we have a total of 23 transmission events with 1 mutation in 37markers. The two lines back to Henry Fox, III, go back 7 generations through two sons of Henry Fox III; Thomas of Virginia, who married Elizabeth Hancock, and William of South Carolina, who married Sara Carroll.

263208 now takes the connection back two more generations directly to Henry Fox 1st and Anne West.  He descends from their son Thomas who married Mary Tunstall, through their son Joseph who married Mildred Fenton,The others all descend from son Henry Fox 2nd, who married Mary Kendrick, the parents of Henry Fox III. On 37 markers, 263208 differs from 48275and 99137 only at DYS 460 where he has 10 repeats and they have 11.  With nine generations in play, a match on 36out of 37 markers is highly significant. The three match at CDYa, showing that this mutation probably occurred only in the line down from Henry Fox III’s son Thomas and prior to their most distant common ancestor, Felix Fox of Lexington County, SC, who married Annie Elizabeth Evans in 1878.  We now have the haplotype of Henry Fox 1st defined with the exception of DYS 460.

This result is highly significant since some researchers had questioned that Henry Fox, 2nd, was actually the son of Henry Fox, 1st.  The story is that his sons Thomas Fox and John Fox are mentioned in the will of their maternal uncle John West (son of Col. John West, the brother of Lord De La Warr), whereas Henry Fox 2nd is not.  This could well be because Henry was first in line to inherit his own father’s estate; nevertheless doubt remained in the mind of researcher Ellen Cocke and others. This result is pretty strong confirmation in some minds. The project would like very much to get a male descendant of Thomas Fox (b abt 1680, who married Mary Tunstall) and of John Fox, (b abt 1676) into the project. There are other reasons to believe that Henry Fox 2nd was the son of Henry Fox 1st (i.e. they held similar church and public offices about 25 – 30 years apart) but it would be good to have Y-DNA verification.

We know there are more of these descendants around. The father of Henry Fox 1st, John Fox, married about 1699 and left Fox descendants.  Dr. J. Ben Robinson, one of his descendants, states that his wife probably was a daughter of Edmund Bacon of New Kent County.  In addition, there was a genealogical group in Washington, DC, back in the late 1930's called"The Society of the Descendants of the Hon. Henry Fox and Anne West” under the leadership of George H. S. King, a Thomas Fox descendant.  Henry Fox 1st, is also said to have had a brother named John Fox b abt 1652, who married a Lightfoot (either Margaret or Frances) and left descendants in Virginia and Maryland.  Hopefully some member of the Virginia Historical Society will read this and take note.

48275 has been SNP tested by Ethnoancestry and is S21+ andS29-.  He has also been tested L47+ by FTDNA.  This is a brand new SNP dividingL48, which in turn divides S21 (S21 is called U106 by FTDNA.)  He would also be S26-, of course.  (In FTDNA terms, he is U106+, L48+, L47+,U198- and L1-).  FTDNA has now recognizedR-L47 in their Haplotree.   ISOGG calls HaplogroupR-47 by the long name R1b1a2a1a1a3b2a.

These Henry Fox descendants match 35 for 37 and 63 for 67 with 26383, a descendant of George Elder, born a slave in 1831, in Rutherford County, Tennessee. Deviations are 15 vs 16 at DYS 458, 18 vs 17 at DYS 570 and23 vs 21 at DYS 413a.  108898 and 117867also deviate from 26383 at CDYa, having 35 rather than 36 repeats.  These markers are all noted for having a high mutation rate but these four deviations indicate that the common ancestor would probably have preceded Henry Fox, 1st, in the line of succession (i.e. an as-yet-unidentified American Fox line.)  26383 has also tested Haplogroup R-L47. The exact connection is being pursued but has yet to be established.

A number of new SNPs are now being offered, which divide L47.  L44 and Z159divide L47 and L46, L45 and L164 further divide L44.  The project will monitor progress as these SNPs are tested and, if they appear useful, further testing will be recommended.  Meanwhile, several of these Foxes have joined the L48 Project at FTDNA. Neal Fox is also advocating 67 or 111 marker tests in the future to help him define subclusters and he reports that a 13 at DYS492 (in the 67marker last panel) points toward a possible S21+ status.

Interestingly enough, both thenull439 and the Henry Fox/ Anne West groups have been found belonging to Founder haplotype clusters that include names such as Callaway and Smith.  This leads us to believe they may have both once belonged to a clan, containing both Haplogroups, that may be of Norman origin – Callaway being a Norman name. All this may eventually help to confirm the Vaux/Fox Norman connection, though it is much too soon to draw conclusions.

One attempt to confirm a Vaux/Fox connection (as predicted by Anthropologist John William Fox) was unsuccessful. It is known that there was a name change from de Vaux to de Strickland in about the year 1200.   Project member 40501, a representative Strickland, was tested by FTDNA for the Fox Project.  Already tested by Relative Genetics, he is not a null 439 and is a genetic distance of 12 from the 37 marker null439 modal. He is also a genetic distance of 15 from the Henry Fox/Anne West descendants, the main candidates for the Vaux/Fox connection.

Andrew Fox Descendants

It is known that descendants of Andrew Fox (1750-1819) of Virginia and Tennessee, lived in close proximity to the Elder family in Rutherford County, TN, in 1831 but this possible Fox/Elder connection has now been denied with the acquisition by the project of two known descendants of Andrew Fox.  They are a genetic distance of 23 to 24 from 26383 in 37 markers.

131451 and 131454 are fourth cousins once removed, the most recent common ancestor (MRCA) being Jacob Fox (1785-1830) of Tennessee, who married Elizabeth Ann Broyles. Jacob Fox was the son of Andrew Fox.  There are 11 transmission events separating them and they have a genetic distance of 5 in 37 markers, though only three markers are involved – all fast mutating markers in FTDNA’s 26 through 37 Marker panel. They are estimated by FTDNA to be M-269+ or Haplogroup R1b1b2.

There is also a very close match with 9DW26 in Ysearch, surname Hughes, who indicates a Scottish ancestry and has tested P312+ and M222-.  The comparison is as follows:

     131451 is 20 at DYS 576, 19 at DYS 570 and 38 at CDYb
     131454 is 18 at DYS 576, 18 at DYS 570 and 40 at CDYb
     9DW26 is 19 at DYS 576, 18 at DYS 570 and 37 at CDYb

Their closest match in the Fox Project (GD=16 or 17 at 37 markers) is 58674, who has been tested P312+ and L21+ (Haplogroup R1b1b2a1b4c). Our guess is that they would all test L21+. Their ancestry is thus projected to be ancient Scots-Irish.

Choctaw County, Mississippi, Foxes

In 80721, 56980 and 62766 we have three members of a Fox group living in Choctaw County, Mississippi, in the late 1800s.  In the case of 80721, the great grandfather was Thomas J. Fox b 1820 Walton Co GA.  In the case of 56980 and 62766, first cousins, the great grandfather was James Davis Fox, born 1824 in Alabama. A later addition to this family grouping is 121692, a descendant of Henry Fox, born ~1750 in Gloucestershire, England.  In 37 markers, there are only three one-step mutations within the group, at DYS 448, CDYa and GATA H4:

      56980: DYS448 = 18 CDYa = 38 GATA H4 = 12
      62766: DYS448 = 19 CDYa = 38 GATA H4 = 12
      80721: DYS448 = 19 CDYa = 37 GATA H4 = 12
    121692: DYS448 = 19 CDYa = 38 GATA H4 = 11

The haplotype of the MRCA can be inferred by triangulation to be the same as 62766. 

The original hypothesis was that Thomas J and James D. Fox were brothers, sons of Johan Fuchs an immigrant from Perish, Germany.  If brothers, there would have been two mutations in 37 markers in 10 transmission events for a mutation rate of 2/(37*10) = 0.0054. This is higher than normal but still well within the expected range.  The addition of 121692 throws an entirely new light on this group.  Interestingly enough, they are also a GD of 5 in 30 markers with a Fox in the SMGF database, whose ancestor was John Fox, 1827, from Monroe, NY.

80721 and 121692 have both been tested at 67 markers and there are another two mutations at DYS413a and at DYS 534.  They are both 13 at DYS 492, indicative of Haplogroup R-U106/S21.  Neal Fox feels they might also test L47+.
It is known that the above Johan Fuchs had two other sons, Jesse and Daniel, and project member 114418 was recruited by this group to join the project because he is a known Jesse Fox descendant. His Y-DNA test results show that 114418 is from a different family entirely (see below under Haplogroup I.)

William Fox of Loudoun Co., VA, Descendants

58674 had originally proposed a connection to Henry Fox and Anne West through their son Thomas Fox.  The genetic distance of 24 at 67 markers was too great to support this contention.  Like 80721 and 48275, he has that 13 at DYS492 that indicates Haplogroup R1b1c9* (S21+) but was tested by Ethnoancestry and found to be S21-.  Actually, this result was not entirely unexpected as he belongs to a cluster Neal Fox has identified within R-M269 that has this characteristic. He has now been tested by FTDNA and found to be P312+ and L21+, putting him in Neal’s Scots-Irish cluster of Haplogroup R-L21.  He is also in the SMGF database and has a total of 78 markers tested.

A match for project member 58674 was then found in the Relative Genetics database (who now has joined the project as Kit B2043) and the most distant known ancestor has now been identified as William Fox, b. 1710 in Loudoun County, VA. Based on research by Kevin Daniels, it appears that 58674 is descended from William’s son James Fox (~1740, VA) through James’ son John Fox (1780, VA) who married Elizabeth Hoffman and the Relative Genetics Fox is descended from William’s son, William Fox, Jr., (~1742, VA) and William, Jr.’s son James Fox (~1789, VA) who married Margaret Franklin. Both are 7th generation descendants of William Fox, Sr., and they match exactly on 39 markers.  Based on the proposed paper trail they would be 5th cousins.

112106, whose most distant known ancestor is Enos Fox, b 1814 in Kentucky, is GD=1 from 58674 at 37 markers and GD=2 from 58674 at 67 markers. There is obviously a family connection here that needs a paper trail and Kevin Daniels has posited that Enos was the grandson of the above James Fox (~1780, VA) and possibly the son of James’ son Jesse  (bet. 1781-1784, VA.)  He bases this largely on the feeling that the descendants of William Fox, Jr., are too well established to include Enos.

164558, a sixth generation descendant, traces back through William’s son, William Fox, Jr., who married Mary (Polly) Brown in 1774 – as does our Fox tested by Relative Genetics and SMGF. It was hoped that both their results at DYS 448 would match, making this a distinctive marker for identifying descendants of the two sons of William Fox, Sr.  Instead, his result was 20 repeats vs 19 for the other. This must have occurred within the last three or four generations (i.e., since 1818), since the two descendants of William Fox, Jr. are third cousins once removed. A mutation from 19 to 20 repeats also occurred in the Enos Fox line.  Since Enos Fox was born in 1814, it would appear that these were separate parallel mutations.

67 markers have now been posted for 184502, who is descended from James Fox and his first wife, Mary Bartleson. Also 19 at DYS 448, he matches and helps define the modal haplotype for this group – a total of 75 markers tested including the results at Relative Genetics and SMGF. No question but that this is a related family group. The maximum deviation from the modal is only 1 mutation.

58674, 112106 and 184502 have been tested at 67 markers by FTDNA and the only deviation in the last 30 is at DYS 577, where 184502 and 112106 are 16 repeats and 58674 is 15 repeats. Probably, 16 was the ancestral value and 58674 has the mutation.
Based on results for 58674, we know that this group of Foxes are members of Haplogroup R-L21, a subclade of Haplogroup R that, in Great Britain, has a very high representation in very old Scots and Irish ancestry. One subclade of this Haplogroup is R-M222, the group of descendants of the Irish king Ui Neill. 58674 is, however, negative for M222 and other known downstream SNPs so we can call him Haplogroup R1b1b2a1b5* or R-L21*.  “Haplogroup R-L21* - Descendants of William Fox of Loudoun Co., VA”, is now a separate grouping in the Y-DNA Results tables on this Website.

Three more recent additions to this family group are 204318, 210491 and N32693.  They are all descendants of Hugh Fox (1745-1820) either of Virginia or North Carolina.  They are a genetic distance of 3 or 4 on 37 markers from the others, which may point to a more distant relationship, but definitely belong to this group.

John B. Fox of Orange Co., VA, Descendants

Another of our Southern Fox group, 93795 is a match for 26653 at 12 markers.  Both trace back to John B. Fox, b: 1745 in Orange County, VA, who married Ann Barber. The MRCA is their son Henry Fox, b. 1801, and the two are third cousins, once removed.  A typical R-M269 haplotype, 93975 has no other close matches in the project.  Tested at 37 markers, his closest 37-marker match in the Fox Project is 65065 at a GD of 13.

Francis Fox, 1785, Wilkes Co.,NC

59807 and 159919 are descendants of Francis Fox,Jr., born: ~1785 in NC or SC  and died: between 1830-1840 in Wilkes County, NC.  The MRCA is William Moses Fox who married Morning Ayers, thus making them fourth cousins.  William Moses Fox was born: 15 Sep 1804, Wilkes Cty, NC, and died: 25 Apr 1882, Yancey Cty, NC.  The two men match exactly on 37 markers.  Theirs is another typical R-M269 haplotype.  They have now been joined by 236252 who differs from them at 439, 447 and CDYb.  236252 is a descendant of William Fox b 1784, PA, d 1853, OH and the connection back to Pennsylvania appears worth exploring.

Francis Fox, Sr., is believed to have been born in England in 1749.  He died ~1821 in Wilkes Co., NC.

Other Southern USA R1b Foxes

We have yet to find a match for a large group of these Southern R1b Foxes:

24011, tested at 12 markers, is a descendant of William Eires Fox, born 1758 in Virginia.  His results have recently been augmented to 38 markers based on SMGF testing but he still has no close matches within the project.

30540, a descendant of David Fox of SC and TN, has been tested at 67 markers but has no close matches within the project.  His closest match is 93372 with a GD = 18 at 67 markers.  He was originally referred to the project by 24011 because of common geographical location of ancestors.  He is a 13 at DYS 492 indicating possibly Haplogroup R-S21/U106.

38215, by virtue of close geographical proximity, had thought that his ancestor, John Fox (1678-1748), of Essex Co., VA, was the son of John Fox (1652-1703) who married Mary Tunstall, the grandson of Henry Fox and Anne West.  At 67 markers he is GD = 31 from 48275, so this connection is disproven.  His closest match at 67 markers is 28579 (GD=21.)

An ardent genealogist, he now writes, ”My line of descent comes from small intrusive Forz (Fortibus) family of Cockermouth, Cumbria (outside of Whitehaven) and Yorkshire, whose first ancestor was a French Count of Aumale who arrived in 1189 from Brittany. His son, William de Forz, became the Earl of Albemarle and was given lands in England in Norfolk, Cumberland and Lincolnshire.”  He has established that his line became goldsmiths and merchants in London, helping to finance the Virginia Company.

He is 12 at DYS 492 and has been deep clade R SNP tested as Haplogroup R1b1b2a1 (L1- L21- L23+ L48- L49+ M269+ P107- P310+ P311+ P312- U106- U198) and he has joined the ht35 project run by Vincent Vizachero, which covers his subclade. This result is indicative of an Eastern Mediterranean origin and is consistent with research indicating that the Forz family may have come from Milan or nearby Genoa, migrated to France where they appear in the court of Eleanor of Aquitaine, if not before. They held manors as far north as Normandy, France and were the admirals of Aquitaine/Poitou.

Conservatively, we have listed John Fox, b1602, London, England, as his most distant known ancestor.  John Fox, an apprentice goldsmith of London, emigrated to Jamestown, VA, in 1635.

45680, a descendant of Gatus Fox of Tennessee, has been tested at 37 markers with no project matches. FTDNA predicts Haplogroup R-M269.  His closest project member is 50481 with a GD of 15 at 37 markers.

59573, tested at 12 markers is descended from George C. Fox of Amarillo, TX, born before 1898.  He is a typical R-M269 but his closest matches in the project are a GD of 2 at 12 markers.  He has also tested mtDNA and is estimated to be mtDNA Haplogroup T*.

88154, a descendant of William Fox, b 1836 in Warwick Co., VA, is another predicted Haplogroup R-M269. His closest matches (31 for 37) are named Tucker (English) and Pridgeon (French) and Neal Fox reports that they form a distinctive cluster with an ancient founder. Within the Fox Project his closest match (only 23 for 37) is 28579. He has some very distinctive results in the first 12 markers but is close to modal for the rest.

107545, tested at 37 markers, is another of our R-M269 Foxes with Southern US ancestry who has yet to find a match.  His most distant known ancestor was William D. Fox, 1887, Alabama.  He might have had ancestors in the Bahamas.

125352 had a great grandfather named Fuchs who came from Alsace, France, to America in the late 1800s. His 12 marker haplotype indicates Haplogroup R-M269 or one of its subclades.  He has no close matches in the Fox Project.  His mtDNA result (HVR1) indicates Haplogroup HV*.

123131 is the great grandson of Hyram Fox of Hart, Michigan.  Hyram came down to Michigan from Canada and is thought possibly to have had Jewish ancestry.  His is an unusual R1b haplotype – close to the modal for R-M269 but with multi-step mutations at DYS 389i and at DYS 438 – both rather stable markers.  He is also 13 at DYS 492, indicative of U106/S21 positive status.  There are some similar haplotypes originating in Belarus/ Lithuania but he matches closely (GD = 1 in 33 markers) a Webb in the SMGF database whose MDA is given as Robert Webb, b 1755 in Crewkerne, Somerset ENGLAND.  123131 has been tested Haplogroup R-U106*.  

121692 is descended from Henry Fox, b ~1750 in Gloucestershire, England.  He is a close match to 56980, 62776 and 80721 who have traced back to Alabama and Georgia around 1820.  At 37 markers he is GD=1 vs 62776 and GD=2 vs 56980 and 80721.  Several more mutations have shown up in the last 30 markers of the 67 marker set against 80721 and it is more probable that the most distant common ancestor (MRCA) of the combined clan is before rather than after Henry Fox.  This group appears to be Haplogroup R-U106, based on the result of 13 repeats at DYS 492.

Two other Southern US Haplogroup R1b1c Foxes have been identified in the SMGF database and partial results are available.  Ancestors are: Benjamin Fox, VA, 1839 and John Fox, Jr., VA, 1780. As yet, no close matches are found for them in the Fox Project.

Haplogroup R1b - Other R1b

This group includes those in Haplogroup R1b (probably all R1b1c) with Northern US, British or German ancestry plus one from Romania and several who have provided no ancestry.

Descendants of Levi Fox and William Fox of Pennsylvania in 1700s

130621 is a descendant of Levi Fox, Sr., (b. 1802 in Washington Co., Pennsylvania).  He is a 12 marker match with 48443, who is a descendant of Taylor Fox, born October 1846 in Sheffield, Tippecanoe Co., IN.  Further research indicates that Taylor’s father, possibly named Isaac, was the brother of Levi Fox and that their parents may have been Bonham and Temperance Fox, born 1760s in Loudon Co., Virginia. Then there are three unproven earlier generations going back to George, b. 1662 in Leichester, England and Jane (Palmer) Fox, born ca 1670. Upgrading 48443 to 37 markers is strongly recommended.

Two new project members, 164677 and 168178, match the Levi Fox, Sr., descendant on 34 out of 37 markers (GD=3) and 35 out of 37 markers (GD=2,) respectively.  They are third cousins, descending from two sons of John Fox, who was born in 1797 in VA and died in Ohio. Also estimated to be in Haplogroup R-M269, they are a 36 for 37 match, with 164677 being 12 and 169178 being 13 at DYS 442.  There is good evidence from a family bible that the parents of John Fox were, like Levi Fox, Sr., born in Pennsylvania.  Their names were William (b 1777) and Sarah (b 1792) Fox and they both died in 1857 in Ohio.  The common geographical connection (Pennsylvania) should certainly be followed up.

130621 has been tested at 67 markers and is 13 repeats at DYS 492, a very strong indication of Haplogroup R-U106 (R1b1b2a1a).

Haplogroup R1b - Other R1b No Matches in the Fox Project

28579 had originally thought his ancestry traced back to Wiltshire (and a brother of Sir Stephen Fox) but more definitive evidence points to Thomas Fox born about 1608 in Melton Magnum, near Norfolk, England, as the most distant known ancestor. 28579 has now been tested at 67 markers by FTDNA and is a genetic distance of 20 or more with the null439 Foxes. He was originally tested on 17 markers in 2002 by Jobling and King and also has partial results in the SMGF database, giving a total of 72 markers in all.

He had been found by EthnoAncestry to be S28/U152+, as predicted, and has now been deep clade tested by FTDNA and is Haplogroup R1b1b2a1b4c1 or R-L20 (L20+ L21- L4- M126- M153- M160- M269+ M65- P312+ SRY2627- U152+.)  He is a member of a STR haplotype cluster identified by Neal Fox that includes an L20+ named Faux but the connection would probably be before the adoption of surnames.

38430, ancestry Chorley, Lancashire, England, has become the most tested project member: Tested at 67 markers by FTDNA and at 74 STR markers by Relative Genetics, he has also done mtDNAPlus testing (Haplogroup T2) and has also done Autosomal Testing - Panel 1, which may eventually lead to a better understanding of his overall family heritage.  He has also ordered the full mtDNA genome. He is Y-Haplogroup R-P312 (R1b1b2a1b*) according to FTDNA and ISOGG (M173+ M207+ M269+ M343+ P25+ P312+ M126- M153- M160- M18- M222- M37- M65- M73- P107- P66- SRY2627- U106- U152- U198.) He should also test for the L21 SNP.

48088, tested at 12 markers, has also supplied results from Oxford Ancestors on a fourth cousin, once removed, both tracing back to William Fox born 1597 in Nottinghamshire, England. They match at the 10 markers in common.
50481 provides a striking case of what can be done by a combination of DNA testing with resource to original records.  Originally listed as a descendant of Valentine Fox, born 1535 in Fordwich, Kent, England, his DNA results led him to believe otherwise.  At 67 markers, he matches closely a number of Stewarts and Stuarts, including a known illegitimate descendant of Charles II.  Peter Beauclerk-Dewar collected a series of DNA samples from his kinsmen who were direct male line descendents of Charles II as part of the research for his book “ Right Royal Bastards”. Comparisons show that 50481 has a virtually identical profile on 67 markers to Murray de Vere Beauclerk, the current (14th) Duke of St Albans.

After exhaustive research, 50481 has concluded that he is descended from Sarah Fox, born 1778 in Margate, and that Sarah’s ancestry went back to Valentine Fox, born 1535 in Canterbury, and possibly back to as early as 1320 in Canterbury. At the age of 22, in late 1801, she gave birth to an illegitimate son, Robert.  The father apparently was Aubrey de Vere Beauclerk, Earl of Burford, and soon afterwards 6th Duke of St Albans. He can be placed in Margate at the time when Robert was conceived, in around September 1800.  This Robert was our Project Member’s Fox ancestor but, on the basis of his research and the DNA match, 50481 has listed as his most distant known ancestor: Hato - Knight of Dol c 990 - c 1055, Le Mans, France.

Interestingly enough, he has also found that Sarah was later legitimately married to Edward Longley of Sandwich, a mariner and customs officer.  They had four children of their own and brought up Robert who kept the Fox surname.  In 1833 and 1834 the family emigrated to Fort Edward, New York, and later to Erie County, PA.  Robert married a woman named Charlotte and had four children, living in Sandwich, but he disappeared in 1834 and is thought to have died at sea. 

At a genetic distance (GD) of 15 in 67 markers, 50481 is the closest Fox yet tested to the null439 Fox group, though not a null result himself.  His haplotype is also reasonably close to that of two Fox descendants of John Fox (b 1827 in Monroe, NY), found in the SMGF database, where he is a GD of 5 with 28 markers in common. He is a GD of 7 from the R1b1 Atlantic Modal Haplotype (37 markers.)  He also has results in the SMGF database, giving him a total of 77 markers tested.  He has now tested L2+ and L20-, putting him in Haplogroup R1b1b2a1b4c*.

65065 is of Irish origin. He has been tested at 37 markers and has clearly a typical R1b1c haplotype but is unrelated to other project members or the NW Irish modal. Within the Fox Project his closest match (GD=9 at 37 markers) is 28579.  He is a member of the following projects in addition to the Fox Project:  Ireland-Heritage, Ireland mtDNA, R1b and mtDNA T FGS (full genome sequence.) 
70235, ancestry Clermont Co. OH, has tested at 67 markers and his closest project matches are 28579 (GD = 22) and some of the R1b Southern Fox group.  He is 12 at DYS 492, so is estimated to be Haplogroup R1b1c* (S21-.)

92311, surname Bonham, was admitted to the Fox Project on the basis of a suspected relationship with Peter Fox, 1756, of New Jersey.  Close to the modal for R-M269, he has no close matches in the project.  He has been tested as Haplogroup R-L21.

93372 is a descendant of Matthias Fox who died in 1690 in Borg, Saarland, Germany.  He has been tested at 67 markers and is 13 at DYS 492, indicating probable R1b1c9* status (S21+.)  Tradition says that his ancestors were originally British and moved to Germany in the mid 1600s when the depopulated Saar was trying to attract residents.  He fits a Frisian cluster with ancestors located in England and Wales as well as Germany, Belgium and the Netherlands.  The family tradition is not denied and, in fact, is given some support by Y-DNA testing.

146610 is a descendant of William Fox born 1760 in Cape May, NJ 1760.  He has a typical R1b1b2 haplotype but with no close matches in the Fox Project.

156993 is an Australian descendant of John Fox, born in Staffordshire, England and married in 1831 in Lancaster, England.  His great grandfather William Henry Fox, b 1854 in Staffordshire, was the immigrant to Australia. The Australian deep ancestor is Percy August Fox dob: 1886 place: Bundaburg, Queensland Australia.  While a typical R-M269 haplotype, he has a very distinctive value of 21 repeats at DYS390.  Tested at 37 markers his closest matches within the project are at a GD of 14.  He has been SNP tested M269+. Other descendants of this family are invited to join the project.

A Genebase result has been obtained from a Fox whose ancestry goes back to York, England, and may include name variations such as Faux.  He is typical R1b1c but has no close matches in the project.

Other 12 marker results in Haplogroup R1b include two Y-DNA transfers from the National Geographic’s Genographic Project. N28347 was SNP tested as Haplogroup R1b1c and his ancestry is German. N32673 has chosen not to disclose his ancestry, as has member 48443.  Member 62724 is also in Haplogroup R-M269 and his ancestry is Romanian.  59573 traces back to George C. Fox of Amarillo, TX, before 1898.

Taken as a group, these results continue to demonstrate the wide divergence in Fox Family DNA haplotypes within the R1b Haplogroup.

Haplogroup I

Interestingly, some significant Fox family connections have been found in Haplogroup I. Haplogroup I results have been segregated into Haplogroups I-M253, I-M223, I-M39 and Other Haplogroup I, on the basis of differences in their haplotypes, using the Athey Haplogroup predictor and the tables of Ken Nordtvedt, an expert on Haplogroup I. Enough SNP testing has been done (and is continuing) that we have reasonable backup for these categories.

Haplogroup I-M253 or I1:

This very common Haplogroup is defined by a number of equivalent SNPs: M253, M307, M450, P30 and P40, S62, S63, S64, S65, S66, S107, S108, S109, S110 and S111. It is called Haplogroup I1 in the 2010 ISOGG and YCC Haplogroup trees. <A HREF=" "><strong>Nordtvedt</strong></A> has divided this group into 28 clusters, the main distinctions being Anglo-Saxon, Norse and Ultra Norse.  There are also some Eastern European clusters. The Fox project Haplogroup I-M253 contingent now consists of 15 project members plus 4 more Foxes found in the SMGF database, one from Ethnoancestry and one from Relative Genetics (now One of the project members also has results from SMGF. Origin is both British and German. There are a number of close matches in this group.

Thomas Fox of Concord, MA, Descendants

We now have five Foxes in the project and three more from the SMGF database who trace back to Thomas Fox (1619-1658), who came to Concord, MA, from England in the early 1600s.  69617, N27705 and 142427 all trace back to Thomas' son Eliphalet and their most recent common ancestor is Eliphalet's son Nathaniel (1682-1765.)  69617 and N27705  are 6th cousins once removed, while 69617 and 142427 are 6th cousins in the direct Fox line.  92618 is descended from Thomas' son Samuel and is an 11th generation descendant of Thomas Fox.  70474 has not given his pedigree except to say he is descended from Thomas Fox.
In 37 markers, 69617, 70474, 92618 and 142427 differ only at CDYb, where 69617 and 142427 have 39 repeats, 70474 has 40 repeats and 92618 has 41 repeats.  Their 12 marker results match those of N27705. 

At the 28 markers in common, their 37 marker results duplicate those of the three Foxes in the SMGF database.  Two of these SMGF Foxes descend from Thomas' son Isaac and are 3rd cousins.  The third, like 92618, descends from Thomas' son Samuel and they appear to be seventh cousins, the most recent common ancestor being Samuel's son Isaac.  All three SMGF pedigrees list Samuel Fox (b abt 1593 of Enfield, Middlesex County, England) as the father of Thomas Fox of Concord.

Another project member, 125558, is a known descendant of Jedidiah Lyman Fox, born in New York State between 1831 and 1840, who married Nancy Ann Armfield in Wisconsin. He died about 1916, presumably in Wisconsin.  Thomas Fox did have several known descendants named Jedidiah Fox, b. abt in Norwich, CT,  and his son Jedidiah, b abt 1725.  This line is also through Thomas Fox’s son Samuel and 125558 and 92618 could well be 7th cousins.  It is known that some of this family did move from Connecticut to upper New York State.

Markers that differ (in 37 markers tested) from known Thomas Fox descendants are:

      69617 is 21 at DYS 570 and 39 at CDYb
      70474 is 21 at DYS 570 and 40 at CDYb
      92618 is 21 at DYS 570 and 41 at CDYb
    125558 is 22 at DYS 570 and 39 at CDYb
    142427 is 22 at DYS 570 and 39 at CDYb

92618 and 142427 have both been tested at 67 markers and are an exact match on the last 30 markers.  This Fox line fits closely into Knordvedt's Ultra Norse cluster, indicating Viking ancestry and research by 92618 has indicated that Thomas Fox belonged to St. Olaves, a Norwegian Church in Southwark, London.  In confirmation, both 92618 and 125558 have tested I-P109, this being a subclade of I-M253 indicative of Scandinavian ancestry ( Haplogroup I1c according to FTDNA; Haplogroup I1d1 according to ISOGG.)

N34591 is a 12 for 12 match with the above descendants of Thomas Fox of Concord, MA. Hopefully he will upgrade to 37 or more markers.  His most distant known ancestor was Sylvestor Fox, born in Wisconsin and married to Mary Whipple.  Sylvestor died on February 6, 1865, during the Civil War, in Louisville, Kentucky.

Interestingly, enough, 78547, a descendant of Louis LeRoy, b abt 1610 in Dieppe, Rouen, France, is a 33 for 37 match, though there are multistep deviations at CDYb.  This member actually joined the Fox project for his mtDNA result because his most distant known female line ancestor was Mary Fox b. Jan 24, 1886, in Rosscommon, Ireland.  There is no good reason to suspect a common male ancestor with the Thomas Fox descendants within the last 450 years but there may well be a connection before that time.

178620 and 235626 are more recent additions to this group. The former descends from Thomas Fox through his son, Isaac, and his primary distinction from the others is a mutation at DYS 19.  235626 traces back to Peter Fox b1796 in Gloucester, NJ.  His main distinction from the others is a mutation at DYS 442.

This group also matches closely with two men named Calkins and Caulkins and a Fox connection is strongly expected.

Elijah Fox Descendants

Six matching Foxes in Haplogroup I1 appear to trace back to North Carolina and Virginia. It appears that this clan may have originated in Virginia, went through North Carolina and then through Cocke County, Tennessee, on their way west.  Four of them trace back to Elijah Fox, born about 1775 in North Carolina.  Elijah apparently moved to Cocke County, Tennessee, and fathered three sons, Ransom, Absolom and John Fox.

36288 is a John Fox descendant, 31167 and 56554 are Absolom Fox descendants and 164277 is a Ransom Fox descendant.  36288 and 56554 match exactly at 37 markers.  31167 and 164277 (tested at only 12 markers) match exactly at 12 markers but differ from the other two at DYS 385b, where they show 15 repeats as compared to 14 for the other two.

Two other Fox Project members appear to belong to this family group, though the exact connection has yet to be determined.  76361 traces back to Allen Fox of Wilkes Co., North Carolina, b.1798, and 147651 traces back to James Fox of Virginia, b.1754 (via his son Enoch Fox, b.1774.)  It is possible that Allen Fox was another son of James.

A recent addition, 246191, also descends from Allen Fox and is an exact match with 76361 on 37 markers.  He has been tested Haplogroup I=P109.

76361, tested at 67 markers, matches 36288 and 56554 exactly on 37 markers.  147651, tested on 37 markers, differs only at DYS 477 where he is 21 repeats as compared to 22 repeats for the other three.  Thus the 37 marker ancestral haplotype is well determined and it would appear that a parallel mutation has occurred at DYS 385b in the two 12 marker results.  This is not a truly rare occurrence but it is recommended that both 12 marker results be upgraded to 37 markers. 

All are estimated to be in Haplogroup I-M253 but none have had this confirmed by SNP testing.  The best fit to Nordtvedt’s Haplogroup I clusters is the Anglo-Saxon version of I1a.
They have been joined by a Burris (N30820) who matches them closely (GD=2 in 37 markers) and has been allowed to join the project. Again, these project members did not previously know each other and are still trying to figure out the paper trail connecting them.

Jacob Fuchs Descendants

24157 and 47889, in Haplogroup I-M253, are fourth cousins twice removed, the most recent common ancestor being David Theobald Fox (1738-1823) of North Carolina.  They are an exact match at 25 markers.  David was the son of Jacob Fuchs (1703-1783), who immigrated from Germany (Alsace or Palatine?) to Bucks County, PA, in 1739.

86766 is a GD = 1 from 24157 at 37 markers.  24157 reports that 86766 is his third cousin with Josiah Burel Fox (1830-1900) of North Carolina and Missouri as the common ancestor.  Josiah was a 4th generation descendant of Jacob Fuchs of Germany and Pennsylvania.  Haplogroup I1a has been confirmed for 86766 by SNP testing (M253+.) There has been no deep clade testing but they appear to fall into Nordtvedt’s M-253 Anglo-Saxon clusters.

143344 is a fourth generation descendant of Nicholas Fox, b. abt 1788 in Chatham County NC, buried in Arch Small Cemetery, Ekin, Tipton Co IN, d. 1847 to 1849 in Tipton County IN.  Nicholas was married to Levina Holder.  143344 and 86766 are an exact match at 37 markers, confirming a suspected relationship.

Other I-M253 Results

24106 is a 6th generation descendant of Abraham Fox, b before 1890 in Adair County, Kentucky and moved to Polk County, Tennessee.  His is a typical Haplogroup I-M53 result and some of his closest 37 marker matches within the Fox Project are the Thomas Fox of Concord MA, descendants at a GD of 9 or 10.  He is a GD of 8 from the Roy descendant, 78547.

99981 is another Haplogroup I-M253 but does not match closely the others in this group.  Tested at 67 markers, his most distant known ancestor is Herbert Fox, 1880, Norfolk, England, but one researcher thought there might be a family connection with the Quaker founder, George Fox.  George Fox had no offspring but may have had brothers or cousins who left descendants. His father, Christopher Fox, was a weaver from Fenny Drayton, about 25 miles south of Leicester, England.

Haplogroup I-M223 or I2b1:

This Haplogroup defined by the SNP markers M223 and S24. The latest ISOGG and YCC Trees have this as Haplogroup I2b1.  It is common in England and on the Continent.  Wilhelm Gabert has associated it with the Germanic tribe called the Sugambrer and given it Viking heritage.  Nordtvedt has divided this group into 8 main clusters – Root1, Root2, Root3, Cont1, Cont1a, Cont2a, Cont2b and Cont2c.   Further subclades are I2b1a defined by M284, I2b1b defined by M379, I2b1c defined by P78 and I2b1d defined by P95.

Lancashire Vikings?

31754 and 36120 are both estimated to be Haplogroup I-M223 (Now Haplogroup I2b1, old Haplogroup I1c) and both have ancestors hailing from Lancashire, England.  They are a GD of 6 at 37 markers. 85639, whose ancestor is Johnny Fox of Stokes Co., NC, matches them both with a GD = 4 at 37 markers.  There are mismatches at only seven markers, as follows:

DYS         19        388        439       460        570       CDYb        442         GD from Modal
31754      16         13          12         12         18          40           12                  3
36120      15         14          11         11         18          39           11                  3
85639      15         13          11         11         17          40           12                  1
Modal       15         13          11         11         18          40           12                  0

It is apparent that these men form a cluster and that 85639 is only one mutation from the modal result for this cluster, which appears to match Nordtvedt’s I-M223-Cont.2b very closely. These Lancashire Foxes have been tentatively traced to a group of Vikings who left Dublin, Ireland, to seek refuge in Lancashire in the year 918 AD.

31754 has also been tested by DNA-Fingerprint and has a number of additional markers, markers which continue to support I-M223-Cont.2b.

Descendants of Christian Fuchs of Kutztown

N69127 now has 67 marker results available.  The most distant known ancestor is Christian Fuchs, born 1749, who died in 1814 in Berks County, PA, in Maidencreek, near the town of Kutztown. The family name was changed to Fox from Fuchs in 1825.  It turns out that he is step-nephew to N23128, who matches him at 12 markers.  Aside from that his closest 67 marker match in the Fox Project is 85639 with GD=27.

N23128 has been SNP tested as M170+, M223+ and P19+, putting the two of them in Haplogroup I2b1.  He has not been tested for downstream SNPs below M223.

Haplogroup I-M223  Other Results:

N16652 – upgraded to 67 markers – has given his most distant known ancestor as William Fox born 1791 in Salem County, New Jersey. He also has results from SMGF. He has tested as Haplogroup I-M223* or I2b1*, the asterisk indicating that he is negative for downstream SNPs M284, M379, P78 and P95.  Nordtvedt has predicted that he might test P95+ or Haplogroup I2b1d.

Another member (89347) of Haplogroup I2b1 or I-M223 has now been tested at 67 markers.  He traces back to George Fox who married Mary Wood at Christ Church in Philadelphia, PA, on August 4, 1751.  He is well removed from other members of this Haplogroup I subclade, except for N22211, who is of Irish ancestry.  They are an 11 for 12 match.

133998 gives his immigrant ancestor as Christian Fuchs/Fox, b. 1840 Oberbexbach, Bavaria-d. 1921 Youngstown, Mahoning, Ohio; most distant ancestor Johann Conrad Fuchs, Buergermeister in Dudweiler-Nord around 1766, Kontrolleur of the mine in Burbach, 1776.  His closest matches (GD=13 at 37 markers) are the Lancashire Viking descent (31754, 36120 and 85639).  They are projected to be Haplogroup I2b1* (or I-M223*) but FTDNA projects 133998 to be Haplogroup I2b (or I-P214).  His mtDNA results (HVR1 and HRV2) indicate the very common Haplogroup H.

Haplogroup I-L38 or I2b2

Descendants of Richard Fox and Hannah Williamson

71539 is a documented descendant of Col. Richard Fox (d 1771) and Hannah Williamson of Virginia (see Project Background Section). His relationship with 68387, a descendant of Joaquin Fox, who moved from New Orleans to Mexico in the mid-1800s, has yet to be established. The two match exactly at 37 markers.  68387 has been deep SNP tested by FTDNA and is L38+ L39+ L40+ M161- M170+ M21- M223- M227- M253- M258+ M26- M307- M72- P19+ P214+ P215+ P216+ P217+ P218+ P30- P37.2- P38+ indicating Haplogroup I-L39 (same as I-L38).  This is now called Haplogroup I2b2 by both FTDNA and ISOGG.  He has also been STR tested at 67 markers and has joined the Haplogroup I-L38 Project at FTDNA.  Interestingly enough, this is the Haplogroup of the skeletons found in the Lichtenstein cave, a Bronze Age archaeological site in central Germany associated with artifacts of the Urnfield culture.

Two recent additions are 2068892, uncle of 68387, and 219447, another known descendant of Richard Fox and Hannah Williamson.  He differs from the others only by a mutation at CDYb.

Descendants of Johan Fuchs of Germany

131649 is a descendant of Jesse Fox of Georgia and Arkansas. His great, great, great grandfather was Johan Fuchs of Germany. He is a first cousin of 114418, whose father underwent heavy radiation exposure at Nagasaki in 1943.  They are a GD=1 in 37 markers, the only deviation being at DYS 391 so the radiation did not appreciably skew the comparison. 

The closest match for 114418 in Ysearch is named Brion (Ysearch 8GD73) from Kimburg, France, with whom he is a genetic distance of 5 in 37 markers. He and Brion have 15 at 388, 11 at 454, 11 at 438. Those are unusual values on very slow mutators. The norm is 13, 12, and 10, respectively.  His 9 at 460 is also rare for Haplogroup I-L39. Because they match on some slow markers they may be closer than would otherwise be apparent.  114418 is currently upgrading to 67 markers to check out this situation.  131649 has been tested M170+, meaning backbone Haplogroup I, but we suspect they would both test L38+ and L39+ and both have joined the L-38 Project at FTDNA.  These two are GD=20 at 37 markers from 68367 and 71539.

Other Haplogroup I -  Not Further Identified

N30890, with a 12-marker haplotype, was SNP tested as Haplogroup I but the subclade not determined. His ancestry is German but that is all that is known.

Haplogroup G2

24750, tested at 12 markers, is assigned Haplogroup G2.  He is a descendant of Franz Joseph Fox and his son Gideon Fox, both of whom came from Germany to Pulaski County, Indiana sometime around 1836. 

N28014, also G2, has indicated the Ukraine as his ancestry.

We have found a Fox who appears to be Haplogroup G2 in the SMGF database.  The most distant ancestor listed is Adam Fox of Philadelphia, born before 1800.

Two recent Haplogroup G2 members are 201278 (ancestor is Harry Fox of San Francisco) and 230009  (ancestor is Alexander Maxwell Fox of SC and GA).  No matches in the project

Haplogroup R1a1

Richard Fox and Vaniah Fox of Glastonbury, CT, Descendants

The only match in Haplogroup R1a1 (ISOGG calls this R1a1a), is where 133409, a descendant of Richard Fox, 1641, of Glastonbury, CT, matches 134182, a descendant of Vaniah Fox, who married Abigail Cadwell in Glastonbury, CT, in 1748.  They match on 35 of 37 markers.  A third Fox, N55006, matches the Vaniah Fox descendant exactly at 12 markers.  His MDKA is Harry Francis Fox (1849) of Lydlinch, Dorset.  Unfortunately for the Fox surname, there is a tradition of an illegitimate Fox ancestor of Harry Francis Fox who took his mother’s name, according to one researcher.  This group is actively searching for connections.  See the Project Background section for further information.

N55006 has been tested M198+ and negative for all downstream SNPs making him Haplogroup R1a1* (R-M198*). The others are presumably the same. 

Haplogroup R1a1 – Other Results

46155, tested at 37 markers, has also been assigned Haplogroup R1a1. This is an eastern version of Haplogroup R, which is well represented in European males, including some of Viking descent. The earliest known ancestor of 46155 is Gottlieb Fox (b. Dec 1846, Germany; d. 19 Jul 1903, Belmont County, Ohio.

N57084, a Genographic Project transfer, is another member of Haplogroup R1a1.  Recruited by N16652 and another investigator interested in New Jersey Fox families, he is a direct descendant of Jeremiah Fox, 1829, of Salem, NJ, but traces back to Frederick Fox, ~1728, Palatinate, Germany.

Haplogroup E3b1b1

We have two transfers from the Genographic Project (N19397 and N23138) whose matches in the FTDNA database suggest Haplogroup E3b1b1, found mainly in the Middle East.  Ancestry is unknown.

144536, a descendant of Barnaby Fox born 1750 in Cumberland County, NJ, is also estimated to be Haplogroup E1b1b1.) He is an 11 for 12 match with N23138 but is 10 as compared to 14 repeats at DYS 439.  No matches as yet.

Haplogroup Q

94524 is a descendant of Aron Fuchs of the Ukraine who moved to the United States in 1915 and changed his name to Harry Fox.  He is a confirmed Haplogroup Q and his 25 marker STR haplotype is typical of this group.  A number of his close matches are reportedly of Ashkenazi descent.  He is a 24/25 match in Ysearch with a man from the Ukraine with the surname Lurie.

N34859 is a descendant of Creek Indian Trig Fox.  Only 12 markers but tested Haplogroup Q-L53.

Haplogroup J2 and Subclades

We now have two members of Haplogroup J1 and 6 members of Haplogroup J2 in the Fox Project.  This is a Mediterranean Haplogroup but exists all over Europe to some extent.  The Romans are thought to have brought this Haplogroup to Britain.

In Haplogroup J2:

97877 is a 37 marker upgrade from Genebase.  The Genebase results were matched and, combined, he now has a total of 56 markers tested.  He is estimated to be Haplogroup J2a1-bk (Athey prediction.) His ancestor is Rankin Fox of Staley, NC, b 1881 

110488, a descendant of William Francis Fox, born 1906 in SC was tested at 37 markers and also appears to be Haplogroup J2a1b.  His SNP testing at FTYDNA confirms Haplogroup J2 (M172+).  Testing M67 would confirm J2a1b (FTDNA presently calls this J2f but this is due to change.)

140364 is our first member of the Palatine, NY, Mohawk Valley Fuchs clan and is estimated as Haplogroup J2.  He is a documented descendant of Christoffel Fuchs, born 1608 in Gravenweidt, Germany.  Several of Christoffel’s grandchildren migrated from Niederbieber, Germany, to New York City in 1710 and then migrated to upstate New York.  “The Mohawk Valley Foxes,” a book by Donald P. Fox, documents this family quite well.

153293 is a descendant of Johannes Friederick Fuchs, b 1727 in Germany, who lived in Cumberland County, NJ, in the 1760s and 1770s.   In his book, "The Fox Family From Germany to Southern New Jersey"(1982), Rulon D. Brooks, Sr., states that Frederick Fox and his wife Mary Band emigrated to the US from Germany aboard the ship "Edinburgh" in 5 Sept 1748.  This may or may not be the same Friederick Fuchs.  Johannes Friederick Fox (Fuchs)'s descendents are well documented primarily because they were farmers and stayed in Deerfield Twp, Cumberland Co, NJ, until the present.  This result has also been found to fall under Haplogroup J2* and is a 23 for 25 match with 140364.  Markers 26 through 37, however, include five mismatches. There may well be a Fuchs family connection here in the far distant past back in Germany.

208921 is a descendant of Morris Fuks of Odessa, Russia.  Many matches with other surnames and has joined the WIRTH Project of related Ashkenazi surnames.  Haplogrooup J2a3b1

240688 is a descendant of Samuel Fox of Russia.  Another J2a4b.

In Haplogroup J1:

N51825 is a 12 marker transfer from the Genographic Project who appears to fit Haplogroup J1.  He fits the 6 marker Cohanim modal haplotype but deviates on 3 markers when extended to 12 markers.  He has not given his ancestry but could probably join the Cohen project at FTDNA if he could establish some sort of paper trail.  This haplotype is common among descendants of Aaron, the Jewish Patriarch.

169337 is a descendant of Moishe Fuchs of Kutno, Poland.

Mitochondrial DNA Results

We have a number of mtDNA postings bringing the total up to 31, including 21 men and 10 women. Seven are transfers from the Genographic Project, two of whom prefer to hide their ancestry. Again there are a wide range of results, including mtDNA Haplogroups B2, H, H1c, H5a1, HV*,J1b1, I, K, K1a1b, T*, T1, T2, U2, U4 and U5a1a.  Differences from CRS (Cambridge Reference Sequence) are shown along with the projected mtDNA Haplogroup in the table.  Most are typical European Mitochondrial Haplogroups.  31754, however, is Haplogroup B, which ties in with a Spanish/Indian Mexican descent on the female side.

There is a match at the HVR1 + HVR2 level between 53477 and 56554, who are niece and uncle, respectively. There is also a match between 30540 and 36120 but, in general, matches at this level are not significant genealogically.

mtDNA results for 14179 and 25481 have been identified in the SMGF database.
14179 is Mitosearch MDBCN:
HVR1: 183-, 215G, 224C, 311C, 519C
HVR2: 073G, 152C, 263G, 315.1C, 497T, 524.1C, 524.2A
This is a firm K1a Haplogroup result because of the 497T, the MRKA ancestor being Euphemia Lacy (or Lacey), born in Loudoun County, VA, in ~1786.  23andMe results have extended this to Haplogroup K1a2, based on a result of 11025 C

25481 is Mitosearch DGMRU
HVR1: 183C,189C,193.1C, 356C, 519C
HVR2: 263G, 315.1C
This is a typical Haplotype H result, the most distant ancestor being Frances A. Johnston, born in 1826 in Putnam County, GA.  23andMe has refined this to H1c1

Results from Other Labs:

Results from other labs are listed at the Alternative Fox Surname Project Website.  They include results from Ethnoancestry, DNA-Fingerprint, Relative Genetics, Gene Bank and Sorenson.

The original Fox test data of Jobling and King at Leicester University have now been identified. One of these is a known descendant of Francis Fox, mentioned above, and is classified R1b1c9a. The other (28579) has joined the Fox Project and is listed with Other Haplogroup R1b.  Originally tested at 16 markers, he has now upgraded to a total of 71 markers.

Additional testing at DNA-Fingerprint and Relative Genetics has cast doubt on the results at DYS 464,a,b,c,d for 25481, a member of the null439 Fox group.  FTDNA continues to call it 14,14,16,16 but other results give him 14,16,16,16, which would match the other members of this group exactly at this marker.  It is expected that FTDNA will shortly change their result.

14179 and 25481 both now appear in the SMGF database and have also have results from Relative Genetics (both use the Sorenson Lab for testing)  While they were null at DYS 439, according to FTDNA, the duplicate Y-DNA test run by Relative Genetics, using different primers, actually shows a value of 12 repeats for DYS 439 (the value assumed by FTDNA.)  They have upgraded to 67 markers at FTDNA, who show a genetic difference for them of 5 but this would reduce to 4, using the revised values for DYS 464 mentioned above.  Their full haplotypes include 11 additional Relative Genetics markers not shown in the Y-haplotype tables and these results are identical.

A Fox tested by Genebank (Canada), ancestor Robert Fox of York, England, has supplied his results, which are typical R1b1c.  He and several additional Fox families found in the Sorenson database are listed at the Supplemental Fox Surname Project Website but not shown under Y-Results on this Website.  24157 has had an additional 5 markers tested by Biotix, also not shown here.

We also have information on four Foxes tested recently by (formerly Relative Genetics.)  They have been given RG numbers for identification purposes:

RG6 is probably in Haplogroup I-M223, along with 10 other Foxes, none closely related.  He descends from John Fox (1770-1809) of Cumberland Co., Pennsylvania.  Before this, he suspects Fuchs ancestry from Alsace-Lorraine, Germany. His Ysearch ID is BWUW8.

RG7 is another apparent descendant of Thomas Fox of Concord, MA.  His known most distant ancestor is Martin Fox born in 1785 in New York State. He matches 92618 exactly at the 35 markers they have in common.   Together with the three Thomas Fox descendants found in the SMGF database, he extends the Thomas Fox modal haplotype out to 75 markers and the number of descendants out to eleven.  His Ysearch ID is GQMSG.

RG8 may be a descendant of John Fox of Ontario County, New York, born in 1820 – his great, great grandfather but it is possible his grandfather may have been adopted while living in Williamsport, PA.  He is obviously another member of Haplogroup R-M269 or one of its subclades but has no close matches in the Fox project.  His Ysearch ID is EJ5G5 and he has some interesting close matches with men named Clendenin and Stephenson in the Ysearch database.  Clendenin has Scots descent and may well be Haplogroup R-L21.

RG9 is also Fox Project member 139347.

SNP testing by Ethnoancestry has been mentioned throughout this discussion.  They offer tests on a number of special SNPs that define important subclades of Haplogroups R and I that are not available elsewhere.

No matches have yet been found for other members of this category within the project, but one member (48088) has provided data for a cousin tested by Oxford Ancestors and two others, found in the SMGF database, had John Fox, born 1827 in Monroe, NY, as the common ancestor.  Both of these did match exactly on the markers tested.

Most of our Y-DNA results, along with pedigrees, can be found in the database - go there and search for the surname Fox.

Very early Progress Reports are available at the Fox DNA Project - Family History & Genealogy Message Board