Welcome to the mitochondrial DNA haplogroup U4 project! Haplogroup U4 (Ulrike) is a small Indo-European haplogroup that is particularly prevalent in Finland and Russia. It is found at low frequencies throughout Europe, North America and Asia. We have project members living in the following countries: Australia, Bahamas, Belgium, Brazil, Canada, Czech Republic, Denmark, England, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Lithuania, Macedonia, The Netherlands, New Zealand, Norway, Poland, Romania, Russia, Scotland, Spain, Sweden, Switzerland, Virgin Islands, West Indies, USA.
Description of Ulrike from Bryan Sykes
The clan of Ulrike (German for Mistress of All) is not among the original "Seven Daughters of Eve" clans, but with just under 2% of Europeans among its members, it has a claim to being included among the numerically important clans. Ulrike lived about 18,000 years ago in the cold refuges of the Ukraine at the northern limits of human habitation. Though Ulrike's descendants are nowhere common, the clan is found today mainly in the east and north of Europe with particularly high concentrations in Scandinavia and the Baltic states. (Source: www.oxfordancestors.com/content/view/35/55)
The project is open to everyone who has taken a mitochondrial DNA (mtDNA) test with the Genographic Project, Family Tree DNA or one of the FTDNA affiliates (e.g., IGENEA, DNA Worldwide), and whose results show that they belong to haplogroup U4 or one of its subclades. There is no fee to join the project, and no additional testing is required. If you have tested with the Genographic Project you will first need to transfer your results to the Family Tree DNA database. Simply log in to your Genographic Project page, go to the section "What else can I do with my results", click on "Learn more" and follow the instructions. If you have tested with another company and wish to re-test with Family Tree DNA, please contact the group administrators. A discount is available for those who have previously tested with Ancestry, Oxford Ancestors, DNA Heritage, Genebase or Relative Genetics. The administrators will supply a promotional form on request. The International Society of Genetic Genealogy have a chart on their website showing the differences in the mtDNA tests offered by the various companies which can be seen here
Please provide details of your most distant known ancestor
on your direct maternal line (your mother, your mother's mother, your mother's mother and so on). Click here
to see a chart on the ISOGG website which shows the path of mtDNA transmission. There is also a useful animated video on the SMGF website which can be seen by clicking here
. You can enter the information via the Plot Ancestral Locations menu on your FTDNA personal page. Please provide the full name, year of birth and location of your most distant known ancestor on your direct maternal line. As we are an international project with members from many different countries please avoid the use of local abbreviations (eg, US state abbreviations, UK Chapman county codes) for clarity.
If you have taken the full mitochondrial sequence (FMS) test (also known as the full genomic sequence or FGS test) please consider sharing your coding region results
with the project administrators. The coding region results are not shown on the project page and will not be made public, and by default they are also hidden from the project administrators. The coding region results enable us to identify your lowest level daughter group (subclade). We can often provide a more detailed classification than that provided by FTDNA. Coding region results can also be used to estimate the age of each subclade, to identify new subclades and to fill out the fine details on the U4 tree. To share your coding region results with the project administrators, please log into your FTDNA account, select My Account and click on Results Display Settings. In the section "Show My mtDNA Coding Region Mutations to Administrators of these Projects", make sure that you have indicated "Yes" for the U4 project.
Join the U4 mtDNA Project - in other languages
Currently, we have our "Join Request" in six languages (English, Русский, Deutsch, Français, Español and Italiano). Click here
to see these translations.
We encourage all our members to join the International Society of Genetic Genealogy (ISOGG). It is free to join and all members receive a free monthly e-mail newsletter. You will also be informed of events of interest in your locality. Further information can be found on the ISOGG website
and in the ISOGG Wiki
Useful linksThe U4 haplogroup blog
The Yahoo U4 haplogroup mailing list
Haplogroup U4 group on Facebook
Haplogroup U group on GenealogyWise
Ron Scott's phylogenetic tree for U4a
(not updated since 2011)
Ron Scott's phylogenetic tree for U4b
(not updated since 2011)
Ron Scott's phylogenetic tree for U4c
(not updated since 2011)
Ron Scott's phylogenetic tree for U4d
(not updated since 2011)
The Meaning of Your Mutations
(a custom report by Ann Turner, M.D., for Ron Scott - hg U4b2a1a)
An interactive map showing the mtDNA peopling of the world
- a Community for sharing full mitochondrial sequence DNA results
Roberta Estes' blog post on the mtDNACommunity
PhyloTree's mtDNA global tree
Charles Kerchner's mtDNA log
Ian Logan's haplogroup U4 webpage
mtDNA toolsJames Lick's mtDNA haplogroup analysis tool
Tom Glad's mtDNA Analysis Utility
BLASTING mtDNA Full Genome Sequence (FGS) Results
Scientific papers of particular relevance to haplogroup U4
Melchior L, Lynnerup N, Siegismund HR, Kivisild T, Dissing J (2010). Genetic Diversity among Ancient Nordic Populations. PLoS ONE
5(7): e11898. (Full article available online
Malyarchuk B, Grzybowski T, Derenko M, Perkova M, Vanecek T, Lazur J, Gomolcak P, Tsybovsky I. Mitochondrial DNA phylogeny in Eastern and Western Slavs. Molecular Biology and Evolution
2008 25(8):1651-1658. (Full article available online
Samara Rubinstein, Matthew C. Duuk, Omer Gokcumen, Sergey Zhadanov, Ludmila Osipova, Maggie Cocca, Nishi Mehta, Marina Gubina, Olga Posukh, Theodore G Schurr. Russian Old Believers: genetic consequences of their persecution and exile, as shown by mitochondrial DNA evidence (Report). Human Biology
June 01, 2008. (Report available online
Kristiina Tambets. Dissertationes Biologicae Universitatis Tartuensis. Towards the understanding of post-glacial spread of human mitochondrial DNA haplogroups in Europe and beyond: a phylogeographic approach. Council of the Institute of Molecular and Cell Biology, University of Tartu, Estonia, 2004. (Thesis available online
B. A. Malyarchuk. Differentiation of the Mitochondrial Subhaplogroup U4 in the Populations of Eastern Europe, Ural, and Western Siberia: Implication to the Genetic History of the Uralic Populations. Russian Journal of Genetics
November 2004, Volume 40, Number 11. (Abstract and preview available online
Olga A. Derbeneva, Elena B. Starikovskaya, Douglas C. Wallace, and Rem I. Sukernik. Traces of Early Eurasians in the Mansi of Northwest Siberia Revealed by Mitochondrial DNA Analysis. American Journal of Human Genetics
2002 April; 70(4): 1009–1014. (Full article available online
Other scientific papers with references to U4
Doron M. Behar, Mannis van Oven, Saharon Rosset et al. A "Copernican" Reassessment of the Human Mitochondrial DNA Tree from its Root
. American Journal of Human Genetics
, Volume 90, Issue 4, 6 April 2012, pp675–684. A number of U4 sequences from Family Tree DNA customers were used in this paper.
Helena Malmström, M.Thomas P. Gilbert, Mark G. Thomas et al
. Ancient DNA Reveals Lack of Continuity between Neolithic Hunter-Gatherers and Contemporary Scandinavians. Current Biology
2009 (article in press). (Abstract only available online
.) A discussion can be found on Dienekes' Anthropology Blog
. The article is also discussed on the GenomeWeb website
. Eight U4 samples were included in the study.
B. Bramanti, M. G. Thomas, W. Haak et al.
Genetic Discontinuity Between Local Hunter-Gatherers and Central Europe’s First Farmers. By (3 September 2009) Science [DOI: 10.1126/science.1176869] (Abstract only available online
.) The article is reviewed on Dienekes Anthropology Blog
. Dienekes comments: "Pre-farming populations seem to have been dominated by mtDNA haplogroup U". He quotes from the paper: "it is intriguing to note that 82% of our 22 hunter-gatherer individuals carried clade U (fourteen U5, two U4, and two unspecified U-types". The two U4 samples were from Bad Duerrenberg, Germany (c. 6850 BC) and Spiginas, Lithuania (c. 6350 BC). The paper states that "U4 types show frequencies between 1% and 5% in most parts of Europe, with Western Europe at the lower end of this range, and northeastern Europe and central Asia showing percentages in excess of 7%".
A. Achilli, C. Rengo, V. Battaglia et al
. Saami and Berbers—An Unexpected Mitochondrial DNA Link. The American Journal of Human Genetics
, 2005, Volume 76, Issue 5, Pages 883-886. (Full article available online
O. A. Derbeneva, E. B. Starikovskaya, N. V. Volodko, D. C. Wallace and R. I. Sukernik. Mitochondrial DNA variation in the Kets and Nganasans and its implications for the initial peopling of Northern Eurasia. Russian Journal of Genetics
, Vol. 38, No. 11, 2002, pp.1316–1321. Translated from Genetika
, Vol. 38, No. 11, 2002, pp.1554–1560. (Abstract and preview available online
BooksDNA and Social Networking
, by Debbie Kennett, The History Press, 2011.
The Seven Daughters of Eve
, by Bryan Sykes, 2001. Hardcover: Bantam Press, London, 2001. Paperback : Corgi Books, 2002. US edition: W. W. Norton and Company, New York, 2001.
The Real Eve: Modern Man's Journey Out of Africa
by Stephen Oppenheimer. 2003. Hardcover: Carroll & Graf, New York, 2003. Paperback: Caroll & Graf, 2004.
Genealogical resources:The Female Line: Researching Your Female Ancestors.
By Margaret Ward. Paperback. Newbury, Berkshire, England: Countryside Books, 2003. A step by step guide to researching the female relatives within your family tree. Topics include the problems of name, women in the eyes of the law, and the historical roles of women both in the home and at work.
Who was your mother's mother's mother's mother? Researching matrilineal lines. An online lecture
by Julie Helen Otto, a genealogist with the New England Historic Genealogical Society in America.
We encourage all our project members to join the appropriate geographical project wherever possible. A full list of geographical projects can be found in the ISOGG Wiki
. There is no limit on the number of projects you can join at Family Tree DNA, but admission to a project is subject to approval by the project administrators.
The mtDNA Haplogroup U4 Project is an independent genealogical research study run by volunteer administrators. The project receives no funding, and participants are responsible for the costs of their own tests. The project was organised as a co-operative effort among those who wish to explore genetic testing to advance their knowledge of deep and recent family backgrounds.
Member information and data obtained from the Haplogroup U4 Project must be attributed to the project, administrators, and Family Tree DNA as outlined in the Creative Commons Licence. Please notify administrators when using data for public or private research.
The mtDNA haplogroup U4 project
is licensed under a Creative Commons Attribution-Noncommercial-No Derivative Works 3.0 Unported Licence